Pupinidius Moellendorff, 1901 (subgenus of Buliminus)
Type species: Buliminus pupinidius Moellendorff, 1901; original designation
Pupinidius anocamptus (Mollendorff, 1901) 上曲蛹巢螺/上弯婴螺
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Buliminus (Pupinidius) anocamptus Moellendorff, 1901: 346, pl.14, figs 11, 12; Kobelt, 1902: 850, pl. 121, figs 11–12. Material examined. HBUMM-00332 (eight mature specimens. All measured; specimen-6 dissected): Nanping, (33°12 N, 104°12 E), Sichuan, China, 2004-V-7, coll. M.Wu.
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Type Locality: Nanping (Sichuan, China–previously Gansu, as in the original description). 原甘肃南坪,今四川九寨沟。
(Wu and Zheng 2009) FIGURE 2. A, Pupinidius anocamptus (Moellendorff, 1901), HBUMM-00332–specimen 1.
(Wu and Zheng 2009) FIGURE 9. Pupinidius anocamptus (Moellendorff, 1901), HBUMM-00332-specimen 6. A, general view of genitalia; B, interior view of penis.
Pupinidius chrysalis (Annandale, 1923) 金蛹巢螺
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Buliminus (Pupinidius) chrysalis Annandale, 1923: 391, pl. 16, figs 4, 4a.
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Pupinidius chrysalis Yen, 1942: 254.
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Type locality: Gorge of Mekong River above Yangtse. 长江上游湄公河峡谷
(Wu and Zheng 2009) FIGURE 2. B–D, Pupinidius chrysalis (Annandale, 1923), 3 type specimens of NHM-1923.5.24.1–16 Reg.
Pupinidius gregorii (Mollendorff, 1901) 格氏蛹巢螺/格氏婴螺
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Buliminus (Pupinidius) gregorii Moellendorff, 1901: 345, pl. 14, figs 9, 10; Kobelt, 1902: 849, pl. 121, figs 9, 10.
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Pupinidius gregorii Yen, 1939: 84, pl. 7, fig. 52.
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Type locality: Ho-dshi-kou b. Mu-Kua-tshi (Sichuan).
(Yen 1939) Taf.7, Fig.52. Pupinidius gregorii (Moellendorff), Ho-dshi-gou bei Mu-gua-tshi, Szechwan (Lectotypus SMF-42003), 2 / 1.
(Wu and Zheng 2009) FIGURE 2. E–F, Pupinidius gregorii (Moellendorff, 1901). E, Lectotype, SMF-42003; F, one specimen of paratypes, SMF-42004.
Pupinidius latilabrum (Annandale, 1923) 阔唇蛹巢螺
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Buliminus (Petraeomastus) latilabrum Annandale, 1924: 392, 395, pl. 16, figs 5, 5a.
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Pupinidius gregorii Yen, 1942: 254 (“Pung-tsu-la” mistaken as “Tung-tsu-la”).
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Type locality: West of Pung-tsu-la, Upper Yangtsa (Yunnan).
(Wu and Zheng 2009) FIGURE 3. A–B, Pupinidius latilabrum (Annandale, 1923). A, NHM-Ellis Coln Acc. No. 2136, original series; B, HBUMM-04328-specimen 1.
(Wu and Zheng 2009) FIGURE 10. Pupinidius latilabrum (Annandale, 1923), HBUMM-04328-specimen 1. A, general view of genitalia; B, interior view of penis.
Pupinidius melinostoma melinostoma (Mollendorff, 1901) 灰口蛹巢螺指名亚种/平果口婴螺
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Buliminus (Subzebrinus) melinostoma Moellendorff, 1901: 341, pl. 13, figs 20–22; Wiegmann, 1901: 237, pl. 10, figs 29–33; Kobelt, 1902: 845, pl. 120, figs 20–22.
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Pupinidius melinostoma Yen, 1939: 83, pl. 7, fig. 45.
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Type locality: Shypu am Puiho, River Lum-pu (S Gansu, China).
(Yen 1939) Taf.7, Fig.45. Pupinidius melinostoma (Moellendorff), Shy-pu am Puo-ho, Kansu (Lecotypus SMF-41978), 2 / 1.
(Wu and Zheng 2009) FIGURE 3. C–D, Pupinidius melinostoma melinostoma (Moellendorff, 1901). C, HBUMM-05577- specimen 6; D, Lectotype, SMF-41978.
(Wu and Zheng 2009) FIGURE 11. Pupinidius melinostoma melinostoma (Moellendorff, 1901), HBUMM-05577-specimen 1. A, general view of genitalia; B, interior view of penis; C, interior view of penis, showing V-shaped pilaster and its distal papilla.
Pupinidius nanpingensis nanpingensis (Mollendorff, 1901) 南坪蛹巢螺指名亚种/南坪婴螺
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Buliminus (Pupinidus) nanpingensis Moellendorff, 1901: 346, pl. 14, figs 13–17; Wiegmann, 1901: 249, pl. 11, figs 45–48; Kobelt, 1902: 850, pl. 121, figs 11, 12.
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Pupinidius nanpingensis nanpingensis Yen, 1939: 84, pl. 8, fig. 1.
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Type locality: Nanping (Sichuan, China). 南坪,今四川九寨沟
(Yen 1939) Taf.8, Fig.1. Pupinidius nanpingensis nanpingensis (Moellendorff), Nan-ping, Kansu (Holotype SMF-41994), 2 / 1.
(Wu and Zheng 2009) FIGURE 4. A–D, Pupinidius nanpingensis nanpingensis (Moellendorff, 1901). A–B, respectively HBUMM-05497- specimen 7 & 8; C, Holotype, SMF-41994; D, type specimen, ZI RAS-N193.
(Wu and Zheng 2009) FIGURE 12. Pupinidius nanpingensis nanpingensis (Moellendorff, 1901), HBUMM-05497-specimen 13. A, general view of genitalia; B, interior view of penis.
Pupinidius nanpingensis ambigua (Mollendorff, 1901) 惑南坪蛹巢螺
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Buliminus (Pupinidius) nanpingensis ambigua Moellendorff, 1901: 347, pl. 14, figs 18, 19; Kobelt, 1902: 850. Pupinidius nanpingensis ambigua Yen, 1939: 84, pl. 8, fig. 2.
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Type locality: Nanping (Sichuan). 南坪,今四川九寨沟
(Yen 1939) Taf.8, Fig.2. Pupinidius nanpingensis ambigua (Moellendorff), Nan-ping, Kansu (Lectotypus SMF-41996), 2 / 1.
(Wu and Zheng 2009) FIGURE 3. E–F, Pupinidius nanpingensis ambigua (Moellendorff, 1901). E, Lectotype, SMF-41996; F, one paratype, SMF-41997.
这一种的照片出现在 Yen, 1939: 84, pl. 8, fig. 2 中。而在《中国重要地区陆生贝类》中,作者将出自 Yen 1939 的模式标本似误被分类为“上弯婴螺” (Pupinidius anocamptus) 。
Pupinidius obrutschewi obrutschewi (Sturany, 1900) 奥蛹巢螺指名亚种/欧氏婴螺
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Buliminus obrutschewi Sturany, 1900: 29, pl. 3, figs 26–28.
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Buliminus (Pupinidius) obrutschewi Moellendorff, 1901: 342.
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Buliminus (Pupinidius) obrutschewi eurystoma Moellendorff, 1901: 342, pl. 8, figs 23, 24; Wiegmann, 1901: 246, pl. 10, figs 40–44; Kobelt, 1902: 846, pl. 108, figs 1, 2, pl. 120, figs 23–27. Pupinidius obrutschewi obrutschewi Yen, 1939: 83, pl. 7, fig. 46.
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Type locality: Schy-pa (S Gansu). 甘肃文县石鸡坝镇
(Yen 1939) Taf.7, Fig.46. Pupinidius obrutschewi obrutschewi (Sturany), Liang-dshia-pa, Kansu (SMF-41982, Lectotypus von P. eurystoma Moellendorff), 2 / 1.
(Wu and Zheng 2009) FIGURE 4. E–G, Pupinidius obrutschewi obrutschewi (Moellendorff, 1901). E, type specimen, ZI RAS-N512; F, HBUMM-05672-specimen 12; G, Lectotype, SMF-41982.
(Wu and Zheng 2009) FIGURE 5. A–B, Pupinidius obrutschewi obrutschewi (Moellendorff, 1901), 2 paratypes of SMF-41983.
(Wu and Zheng 2009) FIGURE 13. Pupinidius obrutschewi obrutschewi (Moellendorff, 1901), HBUMM-05672-specimen 1. A, general view of genitalia; B, interior view of penis.
Pupinidius obrutschewi contractus (Mollendorff, 1901) 缩奥蛹巢螺/反向蛹螺
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Buliminus (Pupinidius) obrutschewi contractus Moellendorff, 1901: 343, pl. 13, figs 25, 26, 27; Kobelt, 1902: 847.
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Pupinidius obrutschewi contractus Yen, 1939: 83, pl. 7, fig. 47.
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Type locality: Zw. Li-dshia-pu u. Hsi-ku-tsheng (Gansu).
(Yen 1939) Taf.7, Fig.47. Pupinidius obrutschewi contractus (Moellendorff), zw. Li-dshia-pu und Hsi-gu-tsheng, Kansu (Lecotypus SMF-41981), 2 / 1.
(Wu and Zheng 2009) FIGURE 5. C, HBUMM-04282-specimen 11; D, Lectotype, SMF-41981. E–F, Pupinidius porrectus (Moellendorff, 1901).
(Wu and Zheng 2009) FIGURE 14. Pupinidius obrutschewi contractus (Moellendorff, 1901), HBUMM-04282-specimen 2. A, general view of genitalia; B, interior view of penis, showing cross-sections.
Pupinidius porrectus porrectus (Mollendorff, 1901) 伸蛹巢螺指名亚种
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Buliminus (Pupinidius) porrectus Moellendorff, 1901: 347, pl.14, figs 20, 21, 22; Kobelt, 1902: 851, pl. 121, figs 20–22.
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Pupinidius porrectus Yen, 1939: 84, pl. 8, fig. 3; Yen, 1942: 254 (not Moellendorff, 1902).
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Type locality: Luding County and Wa-sy-kou near Thal des Tung (Sichuan)
(Wu and Zheng 2009) FIGURE 5. E–F, Pupinidius porrectus (Moellendorff, 1901). E, Lectotype, SMF-41998; F, NHM-1902.5.13.20 Reg.
Pupinidius wenxian Wu & Zheng, 2009 文蛹巢螺
- Type locality: 1070 m a.s.l., 32°57 35.5 N, 104°40 41.6 E, foot of Yuxushan hill, Wenxian County, Gansu, China, 2006-IX-27, coll. M.Wu, J.Liu, W.Zheng & L.Gao. 甘肃文县玉虚山
(Wu and Zheng 2009) FIGURE 7. H, Pupinidius wenxian sp. nov., Holotype, HBUMM-05432-specimen 1.
(Wu and Zheng 2009) FIGURE 16. Pupinidius wenxian sp. nov., Holotype, HBUMM-05432-specimen 1. A, general view of genitalia; B, interior view of penis.
Pupinidius pupinella pupinella (Mollendorff, 1901) 豆蛹巢螺指名亚种/偶婴螺
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Buliminus (Pupinidius) pupinella Moellendorff, 1901: 344, pl. 14, figs 1, 2, 3.
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Pupinidius pupinella pupinella Yen, 1939: 84, pl. 7, fig. 49; Yen, 1942: 254 (not Moellendorff, 1902).
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Type locality: Zw. Lidshiapu und Hsikutsheng (SE Gansu).
(Yen 1939) Taf.7, Fig.49. Pupinidius pupinella pupinella (Moellendorff), zw. Li-dshia-pu und Hsi-gu-tsheng, Kansu (lecotypus SMF-41988), 2 / 1.
(Wu and Zheng 2009) FIGURE 6. A–C, Pupinidius pupinella pupinella (Moellendorff, 1901). A, Lectotype, SMF-41988; B–C, type specimens, ZI RAS-N498
Pupinidius pupinella altispirus (Mollendorff, 1901) 高旋豆蛹巢螺/高旋婴螺
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Buliminus (Pupinidius) pupinella altispirus Moellendorff, 1901: 344, pl. 14, figs 4, 5; Kobelt, 1902: 848.
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Pupinidius pupinella altispirus Yen, 1939: 84, pl. 7, fig. 50.
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Type locality: Dangchang. 甘肃宕昌
(Yen 1939) Taf.7, Fig.49. Pupinidius pupinella altispirus (Moellendorff), Tan-tshang, Kansu (lectoypus SMF-41990), 2 / 1.
(Wu and Zheng 2009) FIGURE 6. D–H, Pupinidius pupinella altispirus (Moellendorff, 1901). D, Lectotype, SMF-41990; E–F, type specimens, ZI RAS-N602 ; G, 1 paratype, SMF-41991; H, HBUMM-05723-specimen 2.
(Wu and Zheng 2009) FIGURE 15. Pupinidius pupinella altispirus (Moellendorff, 1901), HBUMM-05723-specimen 14, general view of genitalia.
Pupinidius pupinidius (Mollendorff, 1901) 蛹巢螺/婴婴螺
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Buliminus (Pupinidius) pupinidius Moellendorff, 1901: 345, pl. 11, figs 6–8; Kobelt, 1902: 847, pl. 121, figs 1–5.
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Pupinidius pupinidius Yen, 1939: 84, pl. 7, fig. 51; Schileyko, 1998: 190, fig. 234.
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Type locality: Dshiedshou (Gansu). 舟曲?
(Yen 1939) Taf.7, Fig.49. Pupinidius pupinidius (Moellendorff), Dshie-dshou, Kansu (lectoypus SMF-41992), 2 / 1.
(Wu and Zheng 2009) FIGURE 7. A–E, Pupinidius pupinidius (Moellendorff, 1901). A, Lectotype, SMF-41992; B, 1 paratype, SMF-41993; C–E, type specimens, ZI RAS-N613.
Pupinidius streptaxis (Mollendorff, 1901) 扭轴蛹巢螺/弯曲婴螺
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Buliminus (Pupinidius) streptaxis Moellendorff, 1901: pl. 8, figs 28, 29; Wiegmann, 1901: 242, pl. 10, figs 34–39; Kobelt, 1902: 847, pl. 120, figs 28, 29.
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Pupinidius streptaxis Yen, 1939: 83, pl. 7, fig. 48.
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Type locality: Near Kuang-ting, between Li-dshia-pu and Hsi-gu-tsheng.
(Yen 1939) Taf.7, Fig.48. Pupinidius streptaxis (Moellendorff), Guang-ting, Kansu (lectoypus SMF-41986), 2 / 1.
(Wu and Zheng 2009) FIGURE 7. F–G, Pupinidius streptaxis (Moellendorff, 1901). F, type specimen, ZI RAS; G, Lectotype, SMF-41986.
Pupinidius melinostoma subcylindricus (Mollendorff, 1901) 近柱灰口蛹巢螺
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Buliminus (Subzebrinus) melinostoma subcylindricus Moellendorff, 1901: 341.
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Type locality: Wen-hsien (Gansu). 甘肃文县
There’s no figures of type materials in Wu and Zheng 2009.
这一种在 Wu and Zheng 2009 的论文中没有模式标本的图片,可能需要查阅最新版《中国动物志: 艾纳螺总科》。
Pupinidius porrectus pygmaea (Blume, 1925) 侏伸蛹巢螺
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Ena porrecta var. pygmaea Blume, 1925, Arch. f. Mollusk. Bd. LVII, p. 17; Yen, 1938, 451, Plate I, fig. 6.
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Type locality: Wa-sze-kou (=Wa-sy-kou, Sichuan).
There’s no figures of type materials in Wu and Zheng 2009.
这一种在 Wu and Zheng 2009 的论文中没有模式标本的图片,可能需要查阅最新版《中国动物志: 艾纳螺总科》。
References
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Yen, T. C. 1939. Die chinesischen Land- und Süßwasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft.
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Wang H. J. (王洪建) and Chen D. N. (陈德牛) 2011. 中国重要地区陆生贝类. 甘肃科学技术出版社, 兰州.
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Wu, M., and W. Zheng. 2009. A Review Of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), With The Description Of A New Species. Zootaxa: 1-31.
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吴岷.中国动物志无脊椎动物第五十八卷艾纳螺总科. 2018. 科学出版社